BackEcology: Climate, Life Histories, Populations, Behavior, and Communities – Study Guide
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Seasons, Climate, and Biomes
Earth’s Tilt, Orbit, and the Causes of Seasons
Earth’s tilt (23.5°) and its orbit around the Sun cause different regions to receive varying amounts of sunlight throughout the year, resulting in seasons.
When a hemisphere is tilted toward the Sun, it experiences summer (more direct sunlight, longer days); when tilted away, it experiences winter (less direct sunlight, shorter days).
Equinoxes occur when both hemispheres receive equal sunlight (spring and fall).
Example: In June, the Northern Hemisphere is tilted toward the Sun (summer), while the Southern Hemisphere is tilted away (winter).
Global Circulation Patterns and the Coriolis Effect
The Sun heats Earth unevenly (more at the equator, less at the poles), causing warm air to rise at the equator and move toward the poles.
This creates global circulation cells (Hadley, Ferrel, Polar cells) that drive major wind patterns.
The Coriolis effect is caused by Earth’s rotation, making winds and ocean currents curve to the right in the Northern Hemisphere and to the left in the Southern Hemisphere.
Prevailing winds: Trade winds (east to west near equator), westerlies (west to east in mid-latitudes), polar easterlies (east to west near poles).
Ocean currents are also shaped by wind, Earth’s rotation, and continental positions.
Geography, Climate, and Biomes
Latitude and elevation influence temperature and precipitation.
Rain shadows: Mountains block moist air, causing dry conditions on the leeward side.
Biomes are major ecological communities defined by climate and dominant vegetation.
Major terrestrial biomes: Tropical rainforest, desert, savanna, temperate forest, grassland, boreal forest (taiga), tundra, chaparral.
Example: Deserts occur at 30° latitude due to descending dry air from Hadley cells.
Species Distribution and Adaptations
Climate (temperature, precipitation), latitude, elevation, and geography determine where species and biomes are found.
Organisms have adaptations (e.g., water conservation in desert plants, antifreeze proteins in tundra animals) suited to their biome.
Limiting factors: Temperature, moisture, food, competition, predation, disease, and dispersal ability restrict species’ geographic ranges.
Within a range, populations may be clumped (resources patchy), evenly spaced (territoriality), or randomly distributed (rare in nature).
Life History and Fitness
Life History Traits and Their Importance
Life history traits include age at first reproduction, number and size of offspring, lifespan, reproductive frequency, and parental care.
These traits affect an individual’s fitness—the ability to survive and pass genes to the next generation.
Trade-offs: Limited resources mean investing in one trait (e.g., many offspring) often reduces investment in another (e.g., parental care).
Life History Strategies: r-Selected vs. K-Selected Species
r-selected species: Rapid reproduction, many small offspring, little parental care, short lifespan. Favored in unpredictable or disturbed environments.
K-selected species: Slow reproduction, few large offspring, high parental care, long lifespan. Favored in stable, crowded environments.
Example: Mice (r-selected) vs. elephants (K-selected).
Population Growth and Life History
Birth rates, death rates, age at reproduction, and survivorship shape population size and age structure.
Life history traits influence how quickly populations can grow or recover from declines.
Population Ecology
Population Statistics and Life Tables
Population size (N): Number of individuals in a population.
Birth rate (b), death rate (d), survivorship (lx), fecundity (mx): Key demographic parameters.
Life tables summarize survival and reproduction by age class.
Survivorship curves:
Type I: High survivorship until old age (e.g., humans).
Type II: Constant survivorship (e.g., birds).
Type III: Low early survivorship, high later (e.g., oysters).
Key Population Parameters and Equations
Net reproductive rate (R0): Average number of offspring per individual per generation.
If , population increases; if , stable; if , decreases.
Equation:
Generation time (T): Average age at which individuals reproduce.
Equation:
Per capita growth rate (r): Rate of population increase per individual.
Equation:
Population Growth Models
Exponential growth (density-independent): Population grows without limits.
Discrete:
Continuous:
Logistic growth (density-dependent): Growth slows as population nears carrying capacity (K).
Equation:
Carrying capacity (K): Maximum population size the environment can support.
Density-independent factors: Affect growth regardless of population size (e.g., weather).
Density-dependent factors: Effects increase with population size (e.g., competition, disease).
Population Growth Curves
Exponential curve: J-shaped, rapid increase.
Logistic curve: S-shaped, levels off at K.
Behavioral Ecology
Proximate and Ultimate Causes of Behavior
Proximate explanations: How a behavior occurs (mechanism, development).
Ultimate explanations: Why a behavior evolved (adaptive value, evolutionary history).
Example: Bird song—proximate: hormonal triggers; ultimate: attracts mates, increases fitness.
Types of Animal Behavior
Foraging, mating, parental care, communication, territoriality, cooperation, aggression are major behavior types studied in ecology.
Cost-Benefit Analysis in Behavior
Behaviors evolve when benefits (e.g., increased survival or reproduction) outweigh costs (e.g., energy, risk).
Costs and benefits can depend on individual condition or environment.
Studying Animal Behavior
Experiments can test both proximate (mechanism) and ultimate (evolutionary) explanations.
Behavioral ecologists ask how animals find food, choose mates, avoid predators, communicate, cooperate, and care for offspring.
Kin Selection and Altruism
Direct fitness: Individual’s own reproduction.
Indirect fitness: Additional reproduction by relatives due to individual’s help.
Inclusive fitness: Direct + indirect fitness.
Kin selection: Natural selection favoring behaviors that help relatives reproduce.
Hamilton’s Rule: Altruism evolves if (r = relatedness, B = benefit to recipient, C = cost to actor).
Coefficient of relatedness (r): Probability two individuals share an allele due to common ancestry.
Relatedness influences likelihood of helping, cooperating, or sacrificing for others.
Community Ecology
Ecological Communities and Species Interactions
Community: Multiple species interacting in the same area.
Major interactions: Competition, predation, herbivory, parasitism, mutualism, commensalism.
Interactions affect survival, reproduction, population size, evolution, and diversity.
Competition and Niches
Competitive exclusion principle: Two species using the same limiting resource cannot coexist indefinitely.
Fundamental niche: Full range of resources a species could use.
Realized niche: Actual resources used, limited by competition.
Niche differentiation/resource partitioning: Species use resources differently to reduce competition.
Character displacement: Evolutionary change in traits that reduces competition.
Defense Strategies
Venomous: Delivers toxin via bite or sting.
Poisonous: Harmful when eaten, touched, or absorbed.
Constitutive defenses: Always present (e.g., thorns, toxins).
Inducible defenses: Produced in response to threat (e.g., chemical release after herbivory).
Different defenses evolve based on cost-benefit trade-offs.
Keystone Species and Community Structure
Keystone species: Have a large effect on community structure despite low abundance (e.g., wolves).
Trophic cascades: Changes at one trophic level affect others (e.g., removing predators increases herbivores, decreases plants).
Species Richness and Latitudinal Gradients
Species richness: Number of species in a community.
Generally higher in tropical regions, lower toward poles.
Hypotheses for latitudinal diversity: More stable climate, higher productivity, longer evolutionary time in tropics.
Interaction Type | Effect on Species 1 | Effect on Species 2 | Example |
|---|---|---|---|
Competition | - | - | Lions and hyenas competing for prey |
Predation | + | - | Wolf eating a deer |
Herbivory | + | - | Caterpillar eating a leaf |
Parasitism | + | - | Tapeworm in a mammal |
Mutualism | + | + | Bees pollinating flowers |
Commensalism | + | 0 | Barnacles on a whale |
Additional info: Some explanations and examples were expanded for clarity and completeness based on standard ecology textbooks.